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In this Handbook, Laith Al-Shawaf and Todd K. Shackelford have gathered a group of leading scholars in the field to present a centralized resource for researchers and students wishing to understand emotions from an evolutionary perspective. Experts from a number of different disciplines, including psychology, biology, anthropology, psychiatry, and others, tackle a variety of "how" (proximate) and "why" (ultimate) questions about the function of emotions in humans and nonhuman animals, how emotions work, and their place in human life. Comprehensive and integrative in nature, this Handbook is an essential resource for students and scholars from a diversity of fields wishing to build upon their theoretical and empirical understanding of the emotions.